Long ago, a group of ten Digimon sacrificed themselves to defeat an ultimate evil that threatened the Digital World. These Ancient Warriors created artifacts from their data, the twenty "Spirits", before leaving the Digital World in the care of the three Celestial Digimon. However, one of the three, Cherubimon begins a reign of terror. To counter this, the Celestial Digimon Ophanimon summons five children from the human world into the Digital World via the train-like Trailmon. There, these five Digi-Destined find their respective Human Spirits and "Spirit Evolve" into Digimon, the Legendary Warriors. However, while finding their respective Beast Spirits, the five find themselves hunted by five other Legendary Warriors that serve Cherubimon.
After they defeat four of the them (JP destroys one, another loses his Beast Spirit to Koji and is destroyed by Duskmon for being useless, Zoe destroys another and Takuya gets the last one), Koji Minamoto discovers that Duskmon, the fifth evil Legendary Warrior, is really his twin brother Koichi Kimura. With Takuya Kanbara's help, Koji is able to free his brother, and Kouichi restores the Spirits of Darkness to their true forms and joins the team as the true Warrior of Darkness, Löwemon. After they defeat Cherubimon who is killed by Takuya as EmporerGreymon, the DigiDestined learn that the true mastermind was the very evil the Ancient Warriors sealed away within the Digital World's core, Lucemon. So now the six DigiDestined must try to defeat the two Royal Knights before they scan all of the Digital World, sending it to Lucemon to complete his rebirth. Ultimately Takuya and Koji succeed in destroying the Knights, but not before they free Lucemon. After fighting a losing battle, the Spirits of the Legendary Warriors come to life and along with the spirits of Seraphimon, Ophanimon and Cherubimon (who was purified of his evil by Takuya when he killed him) give the children new hope. The children and the spirits evolve together to form Susanoomon and defeat Lucemon. The Spirits ultimately expel the children from them, take on their human warrior forms and destroy the weakened Lucemon forever with a sword. The children return home to live happily ever after.
Takuya Kanbara
Kanbara Takuya (神原拓也)
Leader of the group. Uses the Human Spirit of Flame to become Agunimon (Agnimon) and the Beast Spirit of Flame to become BurningGreymon (Vritramon). He later received the power to assume the forms of Aldamon (Ardhamon) and EmperorGreymon (KaiserGreymon).
Koji Minamoto
Minamoto Kōji (源輝二)
He was quiet and reserved, the "lone wolf." Uses the Human Spirit of Light to become Lobomon (Wolfmon) and the Beast Spirit of Light to become KendoGarurumon (Garmmon). He later received the power to become Beowolfmon (Beowulfmon) and MagnaGarurumon.
J.P. Shibayama
Shibayama Junpei (柴山純平)
The oldest of the group. Uses the Human Spirit of Thunder to become Beetlemon (Blitzmon) and the Beast Spirit of Thunder to become MetalKabuterimon (Bolgmon).
Himi Tomoki (氷見友樹)
The youngest of the group. Uses the Human Spirit of Ice, to become Kumamon (Chakmon) and the Beast Spirit of Ice to become Korikakumon (Blizzarmon).
Zoe Orimoto
Orimoto Izumi (織本泉)
The main female member of this season, and is very independent. Uses the Human Spirit of Wind to become Kazemon (Fairymon) and the Beast Spirit of Wind to become Zephyrmon (Shutumon).
Koichi Kimura
Kimura Kōichi (木村輝一)
Koji's twin brother. He was separated from Koji when they were babies when their parents divorced. Originally lost his memory and became Duskmon/Velgemon, an enemy to the others, but later became purified. Uses the Human Spirit of Darkness to become Löwemon and the Beast Spirit of Darkness to become JägerLöwemon (KaiserLeomon).
